In anticipation of being able to hold in-person meetings again, KNPS would like you to save the date for Wildflower Weekend 2022. The next annual event will be held April 9th -10th 2022 at the Natural Bridge State Resort Park in Slade, Kentucky.
Save the Date! Fall Membership Meeting
Every year, KNPS hosts a fall membership meeting to discuss yearly updates and upcoming activities. This year, we will be hosting a virtual meeting via Zoom on Saturday, October 23rd at 4pm Eastern time. An agenda and full meeting information will be published in the October Lady Slipper. Please stay tuned if you would like to attend!
Delisting of Running Buffalo Clover
By: Heidi Braunreiter, Office of Kentucky Nature Preserves Plant Ecologist
Running Buffalo Clover (Trifolium stoloniferum) is a perennial clover with showy white flowers and leaves divided into three leaflets, and was historically native to several eastern and midwestern U.S. states. Based on numerous references to clovers in historical literature, it is believed to have been abundant across its historical range in pre-settlement times, which included the Bluegrass Region of Kentucky (Campbell et al. 1988). However by 1900, its extent had drastically reduced, and only five populations in total remained (Brooks, 1983). Throughout the 20th century, there were numerous fruitless attempts to relocate populations, which lead scientists to believe the species was extirpated from states where it was historically common. Finally, in 1983 populations were relocated in West Virginia and Indiana, as well as in northern Kentucky in 1987 (Campbell et al. 1988). With so few populations known, the U.S. Fish and Wildlife Service (USFWS) opted to list the species in 1987 as ‘federally endangered,’ because it was at a high risk of extinction (USWFS 2007).
The decline of running buffalo clover is believed to have been caused by a combination of factors: as land was settled, habitat destruction ensued and competition from non-native invasive species became a problem (Campbell et al. 1988). Another factor was the loss of the American Bison on the landscape. Historical observations noted the clover’s preference for sites where American Bison congregated (Bartgis 1985). Bison likely provided a means of fertilizer, seed scarification, seed dispersal, and the periodic disturbance regime necessary for the running buffalo clover’s life cycle (Campbell et al. 1988). The loss of bison from the landscape would have dramatically affected the species’ propagation.
Today, you will find running buffalo clover in areas that have periodical disturbances mimicking bison; for example, along trails where plants get trampled, in old estate lawns or cemeteries that are regularly mowed, and on stream terraces with occasional flooding events. Running buffalo clover also prefers dappled sunlight in a forested setting. If the forest canopy becomes either too closed or too open, running buffalo clover populations tend to decline in number as it does not tolerate full shade or full sun (USFWS 2021).
Over the past two decades, the Office of Kentucky Nature Preserves (OKNP) has conducted monitoring on running buffalo clover populations in Kentucky in order to look at population trends and manage for non-native invasive plants. Over this time, many new county records were discovered through targeted searches of historical estates, cemeteries, and stream restoration efforts in the Bluegrass Region. Several successful propagation and introductions efforts have also occurred over the years, for example, new populations were introduced and are now self-sufficient at Shaker Village at Pleasant Hill in Mercer County and Taylor Fork Ecological Area in Madison County.
In 2019, running buffalo clover was proposed for delisting by the USFWS, as many populations had been discovered since its initial listing over 30 years prior. In response to the 2019 proposed delisting, OKNP conducted a state-wide assessment of our running buffalo clover populations. Despite set-backs from the pandemic, we were able to survey 61 populations, which covered the majority of the known extant populations in Kentucky. Sites not included in the assessment were already determined to be extirpated, or inaccessible. Most populations in Kentucky occur on private lands, and although obtaining access to private lands can be difficult, we have been grateful for the curiosity, interest, and enthusiastic response from many property owners. Overall, population trends in Kentucky show approximately half of the populations as stable or increasing in size while the other half of populations are declining in size or extirpated (Table 1). Figure 1 shows an updated range map of extant running buffalo clover populations in Kentucky.
Table 1. Population trends of running buffalo clover in Kentucky during the 2019-2020 statewide survey.
| Population Trends | Number of Populations |
| Relatively stable | 11 |
| Increased in size | 18 |
| Decreased in size | 21 |
| Extirpated** | 9 |
| Not applicable* | 2 |
| Total | 61 |
**Majority previously believed extirpated; extirpation confirmed
* Previously uncounted; unable to compare trend
A huge success that came out of the 2019-2020 statewide survey was the discovery of the largest running buffalo clover population in Kentucky. On the Mt. Folly Farm in Clark County, a population of over 6,000 plants was counted along the Upper Howards Creek. Other large populations of running buffalo clover in the state top out around 1,000 to 2,000 plants. The significance of this population led to the creation of a new Registered Natural Area with OKNP at Mt. Folly Farm. The clover occurs in the floodplain of the creek in cattle pastures and riparian woods. This site holds potential for future research as it has running buffalo clover in areas with and without a cattle-grazing regime.
As of September 2021, running buffalo clover was delisted by USFWS as they believe there are a sufficient number of populations occurring on publicly owned and managed lands to sustain the species’ viability (USWFS 2021). As of 2020, 175 extant populations occur across its range in Indiana, Kentucky, Missouri, Ohio, Pennsylvania and West Virginia, and biologists in these states discover new populations every year. Within Kentucky, many populations occur on Wildlife Management Areas where stream restoration projects are ongoing. The key to the future success of this plant is the continued management of populations to keep invasive plants at bay and the continued implementation of appropriate disturbance regimes, be that mowing, grazing, or trampling.
If you are interested in adopting a running buffalo clover population in Kentucky to do yearly checkups to remove invasive species, please contact me! heidi.braunreiter@ky.gov.
Figure 1. Extant range of running buffalo clover in Kentucky as of February 2021.
Bibliography
Bartgis, R. 1985. Rediscovery of Trifolium stoloniferum Muhl. ex A. Eaton. Rhodora 87:425–429.
Brooks, R. 1983. Trifolium stoloniferum, Running buffalo clover: Description, distribution, and current status. Rhodora 85:343–354.
Campbell, J.J.N., M. Evans, M.E. Medley, and N.L. Taylor. 1988. Buffalo clovers in Kentucky (Trifolium stoloniferum and Trifolium reflexum) – Historical records, presettlement environment, rediscovery, endangered status, cultivation and chromosome number. Rhodora 90:399–418.
USFWS. 2007. U.S. Fish & Wildlife Service: Running Buffalo Clover (Trifolium stoloniferum) Recovery Plan: First Revision. Department of the Interior, U. S. Fish and Wildlife Service, Great Lakes-Big Rivers Region (Region 3), Fort Snelling, MN.
USFWS. 2021. U.S. Fish & Wildlife Service. Running Buffalo Clover Final Listing Determination Fact Sheet. https://www.fws.gov/midwest/endangered/plants/rbcl/runningb.html. Accessed 2 September 2021.
Pine Creek Barrens Nature Preserve Field Trip
Kentucky Native Plant Society member Alan Abbott will be leading the second KNPS field trip since the start of the pandemic to Pine Creek Barrens Nature Preserve in Shepardsville. Owned by the Nature Conservancy, the 158-acre property was featured in Thomas Barnes’ book, Kentucky’s Last Great Places.
This open woodland with a prairie-like ground cover hosts a diversity of native glade flora. Other natural communities at Pine Creek Barrens include the dry upland woods which surround the glade. On the southwest boundary, scenic Pine Creek flows through a beautiful mesic ravine forest lined with small limestone cliffs.
A number of rare or infrequent plants grow here, and most years, visitors can see Barrens Silky Aster (Symphyotrichum pretense), Great Plains Ladiestresses (Spiranthes magnicamporum), and Stiff Gentian (Gentianella quinquefolia) bloom in October.
At least eight rare and endangered plant species have been identified at the nature preserve, including the globally threatened glade cress (Leavenworthia exigua var. Laciniata), which has adapted to grow in small depressions on the exposed bedrock. This plant is found only in select areas in Bullitt and Jefferson counties.
Several years ago, KNPS adopted the policy of requiring preregistration for all activities (other than Wildflower Weekend) and for giving members the first opportunity to register for those activities. We send out emails to current members for all KNPS activities at least one week before the activity is announced to the general public. Participation in this trip was limited to 15 people and was quickly filled up by members. If you would like to get advance notification of future field trips, become a member of KNPS. If you join now, your membership will not expire until the end of 2022.
First love
By Debra Parrish
August is time for the giants in the gardens to bloom. Of all the natives, these are my favorites and, ironweed was my first.
Your blooms mesmerized me and when I grew you,
your height fascinated me.
Farmers hate you. They can’t eradicate you with poison so,
they mow you down. A mere image of your true self.
You can be considered a bully. Nuisance is used
to describe you.
Not picky where you grow—fields,
pastures, roadsides, and in the urban garden at
the woodland edge. You are welcomed there, not a bully nor a nuisance.
Prevailing winds constantly at your back,
always standing tall.
The other giants with their issues,
requiring support, to stand tall.
Your presence reminds me year after year, how much
I love the colors of your blooms.
Intense in the sun; subdued in the shade.
Such a welcomed color amongst the other giants
in the urban garden.
You started the pursuit of always having your color
while yellow and white also still in bloom; from the beginning
of the garden season to the end; and, that love affair
started more than thirty-five years ago.
Perception—how it defines you.
David Taylor, Forest Service
Tall ironweed, Vernonia gigantea, is a member of the sunflower family, Asteraceae. In Kentucky, plants begin blooming in August, occasionally, late July, with some individuals blooming as late as mid-October. The plant has alternate leaves each up to 10 inches long and 2.5 inches wide. Stems are typically 5-6 feet tall but can reach 9-10 feet in bottomland. The purple flowers are small, only about ¼ inch long, but because 20-25 are clustered together a head, and 50 to several hundred heads occur in one inflorescence, the plant is showy. Many species of small bees and butterflies will feed on the nectar produced by the flowers. A large plant can produce 1,000 or more wind-blown seeds.
Tall ironweed is a clumpy plant producing multiple stems from each root system, which is extensive and tough, firmly anchoring the plant in the ground. A few caterpillars and beetles eat the leaves, but otherwise the plant is relatively resistant to herbivory. Once established in a field, especially if overgrazed, the plant spreads quickly, aided by bare soil patches, and general immunity from grazing. The plant is difficult to eradicate and is generally controlled by mowing. In a garden setting, it is wise to cut the inflorescence off once the flowers are finished blooming if you don’t want them to spread. Dispose of the old flowers in the trash to prevent spreading of seed.
Debra Parrish lives in the heart of downtown and spends a lot of time and expends a great deal of love on her urban native gardens. She frequently offers summer tours for gardening clubs and other native gardening organizations.
KNPS Announces 2021 Fall Research Grants for Students and Citizen Scientists!
Did you know that the Kentucky Native Plant Society offers small grants to help defer the costs of botanical research, inventory and native plant restoration? Since 2012, KNPS has awarded $8,100 to students working in these areas. Another $750 was awarded in prior years. The total number of grants awarded in any given year is based on the number of proposals received, the quality of proposals and available funding. The graph below shows the kinds of projects that have been funded.
The second type of grant is new and is open to anyone. It will fund
1. native plant inventory, or
2. rare and native plant restoration.
Native plant inventory grants are limited to Kentucky locations and successful applicants will receive a maximum of $250. Rare and native plant restoration grants are awarded to applicants working with native Kentucky plants, preferentially those which are globally rare (G1, G2). Successful applicants will be awarded a maximum of $500. All rare and native plant restoration grants require coordination with the Office of Kentucky Nature Preserves (OKNP) and the Kentucky Native Plant Conservation Alliance coordinators prior to application.
A grant proposal must include:
- A current curriculum vitae;
- A proposal (not to exceed two single-spaced typed pages) describing the proposed research and the role the grant would play in the research;
- An itemized budget;
- A letter of recommendation from the applicant’s major professor or project director; and
- Indicate whether the grant request is for a student research grant, a native plant inventory grant or a rare and native plant restoration grant.
- If the grant is for rare and native plant restoration, include a letter of support from OKNP.
Budget items typically funded include travel to research sites and supplies such as herbarium paper and lab consumables. No personnel time will be funded.
All Grant Proposals are due by October 15, 2021.
If you are interested in applying for any of the KNPS grants, visit the Grants page at the KNPS website. If, after reading the grants page, you have any questions, please email them to: grants@knps.org.
Adopt a Rockhouse in the Red River Gorge!
Volunteers Needed for Upcoming Sandstone Rockhouse Monitoring Project to help protect native plants, animals and archeological resources
The Office of Kentucky Nature Preserves and U.S. Forest Service are seeking to partner with citizen scientists to monitor culturally and ecologically significant sandstone rockhouses and cliffline communities that occur in the Red River Gorge Geological Area. These communities contain many rare and endemic plants and animal species, as well as archeological resources that need your help! Volunteers must submit an application for review, and will be notified of selection in the fall of 2021. Training will be provided and volunteers will begin to help monitor and conserve their assigned rockhouse in 2022. Time commitment is estimated to be approximately 5 working days throughout the year, as well as email correspondence and other requirements listed below. Groups or individuals are welcome to apply.
Why are sandstone rockhouses and cliffline communities in the Red River Gorge Geological Area significant?
Ecology: Sandstone rockhouses are cave like recesses in sandstone cliffs formed by erosion. Dry by nature, they are almost completely sheltered from precipitation; getting water from windblown rain, waterfalls from above or ground water seepage through the sandstone. Compared to the surrounding conditions of the mixed mesophytic forest, rockhouses are warmer in the winter, cooler in the summer, and have lower evaporation rates and higher humidity. Soils found in rockhouses are mostly low pH sand with high levels of some nutrients from saltpeter earth and prehistoric human activities. Due to these conditions, rockhouses have their own unique flora and fauna that include several rockhouse endemic species. Sandstone rockhouses are home to the Federally Delisted White-haired Goldenrod, a species that only occurs in the RRGGA.
Cultural Heritage:
The structure of the rockhouses naturally provide protection from precipitation,
making them valuable as archeological sites. Artifacts and plant materials decompose in other conditions, but are well preserved in the dry, nitrate-rich soils of rockhouses.
Humans have been utilizing rockhouses for thousands of years, and leaving traces of their lives behind. Approximately 12,000 years ago, Native peoples used them as encampments, locations for special activities, bivouacs, and burial sites. Within the last several centuries, rockhouses have been used as dwellings, barns, campsites, schoolhouses, saltpeter mines, shelters for whiskey stills and more. In addition, faunal and flora remains found in rockhouses have helped archeologists better understand the development of agriculture in eastern North America.
What is the impact of recreation on clifflines and rockhouses?
Clifflines and rockhouses are extremely sensitive to human impacts, yet these locations have seen a continued surge in recreational use. In the last 10 years, visitation to the Red River Gorge Geological Area has increased by almost 40%. There has been extensive documentation of the area on social media, guidebooks, and outdoor websites that have increased visitation of hikers, campers, photographers, and other outdoorspeople. The RRGGA is also one of the most popular climbing destinations in the Eastern United States, therefore the rising popularity of rock climbing and bouldering across the country has led to an influx of visitors wishing to climb at “The Red.”
Increased and prolonged recreational presence in the rockhouse and cliffline communities can cause habitat loss and degradation, erosion, eradication of species, and loss of cultural resources. The growing recreational use has increased the need to monitor cliffline and rockshelter sites that are known to contain populations of rare plant and animal species, and/or heritage resources. The data from the monitoring will be used to ensure that appropriate mitigations are in place to protect these sensitive habitats and significant cultural sites.
How do I get involved?
First, check the list of basic Volunteer Requirements and see if this program is right for you:
Adopt a Rockshelter Program Volunteers must:
1. Have an interest in conserving and protecting biological and cultural resources.
2. Be willing to complete required training provided by the Office of Kentucky Nature Preserves and U.S. Forest Service. Training is being developed by OKNP, USFS and Dan Doursan, and will be provided to volunteers in 2022.
3. Be able to fulfill the time commitment required by the program. Adopting 1 rockshelter carries a 5-day commitment that will include training and site visits.
4. Be able to hike long distances on uneven terrain along designated and user defined trails.
5. Feel comfortable working with, or being willing to learn, a touchscreen tablet for data collection.
6. Be able to remove invasive species.
If the ARP is something you’d like to be involved with, please email naturepreserves@ky.gov to receive an application. OKNP and USFS staff will review all volunteer applications and contact those who have been admitted.